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In animals, the baring of teeth is often used as a threat or warning display known as a snarl or a sign of submission. For chimpanzees, it can also be a sign of fear. However, not all animal displays of teeth convey negative acts or emotions. For example, Barbary macaques demonstrate an open mouth display as a sign of playfulness, which likely has similar roots and purposes as the human smile. The influence of smiling on others is not necessarily benign. It may take the form of positive reinforcement, possibly for an underhand manipulative.
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